I wanted to update this thread one more time in case someone runs across it and wants to know if it was resolved. A neighbor had a digital manometer that I was able to borrow, and the regulator that I found had a side tap on it for testing pressure while the grill was operating.
Pressure while grill was off = 7" WC
Operating pressure with one burner on high & one burner off = 4.1" WC
Operating pressure with both burners on high = 3.8" WC
Operating pressure with both burners on medium = 4.1" WC
Operating pressure with both burners on low = 4.2" WC
So Larry, going back to your thought on setting it at mid range flow instead of max flow, it looks like it would not swing the measurement too much using one method or the other. I might set the regulator so that it is at 3" when both burners are on high (worse case scenario), and I should still be less than 3.5" if its on medium or low, so definitely in the same ballpark. And yes, I totally realize I'm overthinking all of this, but once I went down the rabbit hole, I had a hard time coming back out.
I may have to tweak it a little bit after I increase my orifice size ever so slightly based on the chart (new burners are 18,500 BTU vs the old ones which were 15,000 BTU each), but I'm getting really close to the 3" WC operating pressure that Ducane recommends. It is bright sunlight today so I'm having a hard time seeing the flame size and quality, so I need to look again tonight. But it looks like the burners that I got with the donor grill are in worse shape than I was expecting, so I might need to make a small investment.
The other good thing that I was able to test is that my system was holding a constant pressure after I shut off the gas supply. When I had checked it last week with some different fittings and the old manifold, it was dropping slowly when I shut off the valve behind the grill, so there must have been a small leak somewhere.